Alignment of mating pairs during spermatophore exchange: (a) individuals aligned head-to-head with non-simultaneous acquisition of spermatophores (note acquired spermatophore within vaginal pouch); (b) simultaneous reciprocal acquisition of spermatophores of individuals aligned head-to-head; (c) individuals aligned head-to-head with simultaneous acquisition of spermatophores (note: spermatophore still within spermatophore chamber): (d) individuals aligned head-to-haptor with simultaneous exchange of spermatophores - Lim, S.L.H. (2002)